:root {
  --bg: #f3efe6;
  --bg-soft: rgba(255, 252, 247, 0.76);
  --panel: rgba(255, 249, 240, 0.86);
  --text: #1e1915;
  --muted: #625448;
  --line: rgba(50, 33, 19, 0.14);
  --accent: #9c4f2d;
  --accent-deep: #5e2915;
  --accent-soft: #d9a06b;
  --shadow: 0 22px 70px rgba(73, 44, 24, 0.14);
}

* {
  box-sizing: border-box;
}

html {
  scroll-behavior: smooth;
}

body {
  margin: 0;
  color: var(--text);
  font-family: "Avenir Next", "Segoe UI", sans-serif;
  background:
    radial-gradient(circle at top left, rgba(222, 182, 136, 0.42), transparent 34%),
    radial-gradient(circle at 85% 15%, rgba(107, 59, 32, 0.18), transparent 24%),
    linear-gradient(180deg, #f7f3eb 0%, #efe6d8 100%);
  min-height: 100vh;
}

img {
  display: block;
  max-width: 100%;
}

a {
  color: inherit;
}

.site-shell {
  width: min(1200px, calc(100% - 32px));
  margin: 0 auto;
  padding: 24px 0 72px;
}

.topbar {
  position: sticky;
  top: 16px;
  z-index: 10;
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 24px;
  padding: 14px 18px;
  backdrop-filter: blur(18px);
  background: rgba(255, 250, 243, 0.72);
  border: 1px solid var(--line);
  border-radius: 999px;
  box-shadow: 0 12px 30px rgba(83, 59, 36, 0.08);
}

.brand {
  display: inline-flex;
  align-items: center;
  gap: 12px;
  text-decoration: none;
}

.brand-mark {
  width: 40px;
  height: 40px;
  display: grid;
  place-items: center;
  border-radius: 50%;
  background: linear-gradient(135deg, var(--accent), var(--accent-soft));
  color: #fff9f2;
  font-weight: 700;
  overflow: hidden;
  flex: 0 0 auto;
}

.brand-mark-photo {
  background: #d7cab7;
  box-shadow: 0 6px 16px rgba(83, 59, 36, 0.16);
}

.brand-mark-photo img {
  width: 100%;
  height: 100%;
  display: block;
  object-fit: cover;
  object-position: center;
}

.brand-text,
.topnav a,
.button,
.eyebrow,
.project-kicker,
.hero-card-label {
  letter-spacing: 0.04em;
  text-transform: uppercase;
}

.brand-text {
  font-size: 0.9rem;
  font-weight: 700;
}

.topnav {
  display: flex;
  flex-wrap: wrap;
  justify-content: flex-end;
  gap: 18px;
}

.topnav a {
  text-decoration: none;
  font-size: 0.78rem;
  color: var(--muted);
}

.hero {
  display: grid;
  grid-template-columns: minmax(0, 1.4fr) minmax(320px, 0.8fr);
  gap: 28px;
  padding: 72px 0 36px;
  align-items: start;
}

.hero-stack {
  display: grid;
  gap: 18px;
}

.profile-card {
  margin: 0;
  border-radius: 34px;
  overflow: hidden;
  border: 1px solid var(--line);
  background: rgba(255, 250, 243, 0.82);
  box-shadow: var(--shadow);
}

.profile-card img {
  width: 100%;
  aspect-ratio: 1 / 1.08;
  object-fit: cover;
  object-position: center center;
}

.eyebrow,
.project-kicker,
.hero-card-label {
  margin: 0 0 14px;
  color: var(--accent-deep);
  font-size: 0.74rem;
  font-weight: 700;
}

.hero h1,
.section-heading h2,
.feature-card h3,
.project-spotlight h3,
.publication-item h3,
.panel h3 {
  font-family: "Iowan Old Style", "Palatino Linotype", "Book Antiqua", Georgia, serif;
}

.hero h1 {
  margin: 0;
  font-size: clamp(1.95rem, 3.8vw, 3.45rem);
  line-height: 1.05;
  max-width: 13ch;
}

.lede {
  max-width: 62ch;
  margin: 24px 0 0;
  font-size: 1.08rem;
  line-height: 1.8;
  color: var(--muted);
}

.hero-actions {
  display: flex;
  flex-wrap: wrap;
  gap: 14px;
  margin-top: 28px;
}

.button {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  min-height: 48px;
  padding: 0 20px;
  border-radius: 999px;
  text-decoration: none;
  font-size: 0.78rem;
  font-weight: 700;
}

.button-primary {
  background: var(--accent-deep);
  color: #fff5ef;
}

.button-secondary {
  border: 1px solid var(--line);
  background: rgba(255, 252, 247, 0.82);
}

.button-award {
  border: 1px solid rgba(156, 79, 45, 0.24);
  background: linear-gradient(135deg, rgba(156, 79, 45, 0.12), rgba(255, 248, 238, 0.92));
  color: var(--accent-deep);
}

.hero-card,
.panel,
.feature-card,
.project-spotlight {
  border: 1px solid var(--line);
  background: var(--panel);
  box-shadow: var(--shadow);
}

.hero-card {
  padding: 26px;
  border-radius: 30px;
}

.hero-card dl {
  margin: 0;
  display: grid;
  gap: 18px;
}

.hero-card dt {
  font-size: 0.8rem;
  color: var(--muted);
  margin-bottom: 6px;
}

.hero-card dd {
  margin: 0;
  font-weight: 600;
  line-height: 1.55;
}

.section {
  padding: 42px 0;
}

.section-grid,
.contact-section {
  position: relative;
}

.panorama-card {
  margin: 0 0 22px;
  border-radius: 34px;
  overflow: hidden;
  border: 1px solid var(--line);
  box-shadow: var(--shadow);
}

.panorama-card img {
  width: 100%;
  aspect-ratio: 16 / 6;
  object-fit: cover;
  object-position: center;
  display: block;
}

.section-heading {
  display: grid;
  gap: 8px;
  margin-bottom: 24px;
}

.section-heading h2 {
  margin: 0;
  font-size: clamp(1.38rem, 2.5vw, 2.08rem);
  line-height: 1.16;
  max-width: 12ch;
}

.about-grid,
.contact-grid {
  display: grid;
  grid-template-columns: repeat(2, minmax(0, 1fr));
  gap: 22px;
}

.panel {
  padding: 24px;
  border-radius: 28px;
}

.panel p,
.feature-card p,
.project-copy p,
.publication-item p,
li {
  color: var(--muted);
  line-height: 1.75;
}

.pdf-badge {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  min-width: 2.2rem;
  margin-left: 0.55rem;
  padding: 0.08rem 0.42rem;
  border: 1px solid rgba(156, 79, 45, 0.28);
  border-radius: 999px;
  background: rgba(156, 79, 45, 0.08);
  color: var(--accent-deep);
  font-size: 0.68rem;
  font-weight: 700;
  letter-spacing: 0.06em;
  line-height: 1.2;
  text-decoration: none;
  vertical-align: middle;
}

.pdf-badge:hover,
.pdf-badge:focus-visible {
  background: rgba(156, 79, 45, 0.15);
  border-color: rgba(156, 79, 45, 0.42);
}

.research-grid {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 22px;
  align-items: stretch;
}

.feature-card {
  display: flex;
  flex-direction: column;
  border-radius: 30px;
  overflow: hidden;
}

.feature-card:not(.visual-card) {
  padding: 24px;
}

.visual-card img {
  width: 100%;
  aspect-ratio: 16 / 10;
  min-height: 240px;
  object-fit: cover;
}

.visual-card-contain {
  background: rgba(255, 251, 245, 0.96);
}

.visual-card-contain img {
  aspect-ratio: 16 / 10;
  object-fit: contain;
  object-position: center;
  background: rgba(255, 251, 245, 0.96);
}

.visual-card-light img {
  padding: 10px 10px 0;
}

.visual-card-genomics img {
  aspect-ratio: 16 / 10;
  object-fit: cover;
  object-position: left bottom;
}

.fieldwork-spotlight .project-visual img {
  object-position: center;
}

.feature-copy {
  display: flex;
  flex: 1;
  flex-direction: column;
  justify-content: flex-start;
  padding: 22px 24px 24px;
}

.feature-card h3,
.publication-item h3,
.panel h3,
.project-spotlight h3 {
  margin: 0 0 12px;
  font-size: 1.12rem;
  line-height: 1.24;
}

.publication-list {
  display: grid;
  gap: 14px;
}

.publication-item {
  display: grid;
  grid-template-columns: 62px minmax(0, 1fr);
  gap: 16px;
  padding: 22px 24px;
  border-top: 1px solid var(--line);
}

.publication-highlight {
  border: 1px solid rgba(156, 79, 45, 0.18);
  border-radius: 28px;
  background: linear-gradient(135deg, rgba(156, 79, 45, 0.08), rgba(255, 250, 243, 0.82));
  margin-bottom: 14px;
}

.publication-highlight .pub-index {
  font-size: 0.98rem;
  font-weight: 700;
}

.publication-item:first-child {
  border-top: 0;
}

.pub-index {
  font-family: "Iowan Old Style", "Palatino Linotype", "Book Antiqua", Georgia, serif;
  color: var(--accent);
  font-size: 1.1rem;
  font-weight: 700;
}

.project-spotlight {
  display: grid;
  grid-template-columns: 1.05fr 0.95fr;
  gap: 24px;
  padding: 24px;
  border-radius: 34px;
}

.project-visual img {
  width: 100%;
  height: 100%;
  min-height: 280px;
  object-fit: cover;
  border-radius: 24px;
  border: 1px solid var(--line);
}

.gallery-wall {
  column-count: 4;
  column-gap: 12px;
}

.gallery-shot {
  margin: 0 0 12px;
  break-inside: avoid;
  overflow: hidden;
  border-radius: 22px;
  background: rgba(255, 251, 245, 0.92);
  box-shadow: 0 14px 34px rgba(73, 44, 24, 0.12);
}

.gallery-shot img {
  width: 100%;
  display: block;
  object-fit: cover;
  transition: transform 180ms ease;
}

.gallery-shot:hover img,
.gallery-shot:focus-within img {
  transform: scale(1.02);
}

ul {
  margin: 0;
  padding-left: 18px;
}

.icon-link-grid {
  display: grid;
  grid-template-columns: repeat(auto-fit, minmax(180px, 1fr));
  gap: 14px;
}

.icon-link-card {
  display: flex;
  align-items: center;
  gap: 12px;
  min-height: 74px;
  padding: 14px 16px;
  border: 1px solid rgba(50, 33, 19, 0.12);
  border-radius: 20px;
  background: rgba(255, 252, 247, 0.78);
  color: inherit;
  text-decoration: none;
  transition:
    transform 140ms ease,
    box-shadow 140ms ease,
    border-color 140ms ease,
    background 140ms ease;
}

.icon-link-card:hover,
.icon-link-card:focus-visible {
  transform: translateY(-1px);
  border-color: rgba(156, 79, 45, 0.28);
  background: rgba(255, 248, 240, 0.94);
  box-shadow: 0 12px 30px rgba(83, 59, 36, 0.08);
}

.icon-link-mark {
  width: 42px;
  height: 42px;
  flex: 0 0 42px;
  display: grid;
  place-items: center;
  border-radius: 14px;
  font-size: 0.76rem;
  font-weight: 800;
  letter-spacing: 0.08em;
  color: #fffaf4;
}

.icon-link-label {
  font-size: 0.95rem;
  font-weight: 600;
  line-height: 1.35;
  color: var(--text);
}

.icon-link-mark-scholar {
  background: linear-gradient(135deg, #294c7a, #5e8bc1);
}

.icon-link-mark-cv {
  background: linear-gradient(135deg, #465c4f, #7f9a85);
}

.icon-link-mark-award {
  background: linear-gradient(135deg, #8d5b18, #d6a145);
}

.icon-link-mark-lab {
  background: linear-gradient(135deg, #5b2f1b, #9c4f2d);
}

.icon-link-mark-center {
  background: linear-gradient(135deg, #2f5f5b, #5d9d98);
}

.icon-link-mark-reserve {
  background: linear-gradient(135deg, #6a4a1e, #b4873c);
}

.icon-link-mark-media {
  background: linear-gradient(135deg, #7a2f28, #bd5a48);
}

.icon-link-mark-video {
  background: linear-gradient(135deg, #a22d2d, #d94f4f);
}

.icon-link-mark-blog {
  background: linear-gradient(135deg, #35593b, #628d5f);
}

.icon-link-mark-news {
  background: linear-gradient(135deg, #5d364f, #9b6380);
}

.site-footer {
  margin-top: 24px;
  padding: 24px 0 4px;
  border-top: 1px solid rgba(50, 33, 19, 0.12);
}

.site-footer-inner {
  display: grid;
  gap: 8px;
  justify-items: center;
  text-align: center;
}

.site-footer-copy,
.site-footer-meta {
  margin: 0;
  color: rgba(98, 84, 72, 0.92);
  font-size: 0.86rem;
  line-height: 1.7;
}

.site-footer-meta a {
  color: inherit;
  text-decoration: none;
}

.public-security-link {
  display: inline-flex;
  align-items: center;
  gap: 6px;
}

.public-security-icon {
  width: 0.95rem;
  height: 0.95rem;
  flex: 0 0 auto;
  fill: rgba(98, 84, 72, 0.92);
}

.site-footer-meta a:hover,
.site-footer-meta a:focus-visible {
  text-decoration: underline;
}

.footer-divider {
  display: inline-block;
  margin: 0 10px;
  color: rgba(98, 84, 72, 0.56);
}

@media (max-width: 980px) {
  .topbar,
  .hero,
  .about-grid,
  .research-grid,
  .contact-grid,
  .project-spotlight {
    grid-template-columns: 1fr;
  }

  .topbar {
    border-radius: 28px;
  }

  .topnav {
    justify-content: flex-start;
  }

  .hero {
    padding-top: 48px;
  }

  .hero h1 {
    max-width: 15ch;
  }
}

@media (max-width: 680px) {
  .site-shell {
    width: min(100% - 20px, 1200px);
  }

  .topbar {
    position: static;
  }

  .publication-item {
    grid-template-columns: 1fr;
  }

  .gallery-wall {
    column-count: 2;
  }

  .footer-divider {
    display: none;
  }

  .site-footer-meta span,
  .site-footer-meta a {
    display: block;
  }
}